Alward, B.A., Cathers, P.H., Blakkan, D.M., & Fernald, R.D. (2020)
A behavioral logic underlying aggression in an African cichlid fish. Biorxiv.

Recent publications

Alward, B. A., Laud, V., Skalnik, C. J., York, R. A., Juntti, S. A., & Fernald, R. D. (2020). Modular genetic control of social status in a cichlid fish. Proceedings of the National Academy of Sciences, 117, 28167–28174.

Alward, B.A., Hilliard, A.T., York, R.A., Fernald R.D. (2019) Hormonal regulation of social status and temporal organization of behavior in an African cichlid. Hormones and Behavior, 107, 83-95.

Stevenson, T. J., Alward, B.A., Ebling, F. J., Fernald, R. D., Kelly, A., & Ophir, A. G. (2018). The Value of Comparative Animal Research: Krogh’s Principle Facilitates Scientific Discoveries. Policy Insights from the Behavioral and Brain Sciences, 5, 118–125.

Madison, F.N., Kesner, A.J., Alward, B.A., Ball, G.F. (2018) Sex differences in mineralcorticoid and glucocorticoid receptor mRNA expression in response to acute mate pair separation in zebra finches (Taeniopygia guttata). Hippocampus, 28, 698-706.

Alward, B.A.,Balthazart, J.,Cornil, C., Ball, G.F. (2018). The regulation of birdsong by testosterone: Multiple time-scales and multiple sites of action. Hormones and Behavior (Invited Review), 104, 32–40.

Alward, B.A., Balthazart, J., Ball G.F. (2017) Dissociable effects of androgen signaling in cortex-like brain regions on distinct features of birdsong. Journal of Neuroscience, 37, 8612-8624.

For a full list of publications, visit my Google Scholar page. If you can’t download an article, send me an email ( and i’ll be happy to send you a PDF.

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